NEW YORK — Two foreign-born Florida businessmen who helped President Donald Trump's personal lawyer Rudy Giuliani investigate political rival Joe Biden pleaded not guilty on Wednesday to illegally funneling money to a pro-Trump election committee and other politicians.

Ukraine-born Lev Parnas and Belarus-born Igor Fruman entered their not guilty pleas to federal campaign finance charges before US District Judge Paul Oetken in Manhattan.

Federal prosecutors have accused Parnas and Fruman of using a shell company to donate $325,000 to the pro-Trump committee, America First Action, and raising money for former Texas Congressman Pete Sessions as part of an effort to have the Republican president remove the US ambassador to Ukraine.

Prosecutors are examining Giuliani's interactions with Parnas and Fruman, according to a source familiar with the matter.

Giuliani has denied wrongdoing.

Parnas and Fruman's arrest at a Washington-area airport earlier this month was yet another political hazard for Trump amid a fast-moving impeachment inquiry by the Democratic-led US House of Representatives.

The inquiry is centered on the Republican president’s request in a July phone call for Ukrainian President Volodymyr Zelenskiy to investigate former Vice President Biden, a top contender for the 2020 Democratic presidential nomination. The request came after Trump had withheld $391 million in security aid to Ukraine, which he later released.

Democrats have accused Trump of pressuring a vulnerable foreign ally to dig up dirt on a domestic political opponent for his own political benefit.

Giuliani has said Parnas and Fruman helped his efforts in Ukraine to investigate Biden and Biden’s son Hunter. The younger Biden had served as a director of a Ukrainian energy company.

Parnas and Fruman have been asked, as part of the impeachment inquiry, to produce documents and give testimony.

Trump has denied wrongdoing and has described the impeachment probe as a partisan smear.

Prosecutors say Parnas and Fruman committed to raise $20,000 for Sessions as part of an effort to get Trump to remove US Ambassador to Ukraine Marie Yovanovitch.

That effort was carried out at the request of at least one Ukrainian official, prosecutors said.

Trump ordered Yovanovitch removed in May. Yovanovitch testified in the House impeachment inquiry on Oct. 11.

Prosecutors said Parnas and Fruman funneled money from an unnamed Russian businessman to political candidates in several states in order to help the businessman obtain permits needed for a proposed marijuana business, which never came to fruition. US law prohibits foreign donations to political campaigns.

Parnas and Fruman were each charged with two counts of conspiracy, one count of false statements and one of falsification of business records.

Two other men charged in the marijuana business scheme, Andrey Kukushkin and David Correia, pleaded not guilty last Thursday. — Reuters